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1. The spine. 2. The continued stem or midrib of a pinnately compound leaf, as in a rose leaf or a fern. 3. The principal axis in a raceme, spike, panicle, or corymb. 4. The shaft of a feather. The rhachis of the after-shaft, or plumule, is called the hyporhachis. 5. The central cord in the stem of a crinoid. 6. The median part of the radula of a mollusk. 7. A central cord of the ovary of nematodes. 
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